Influence of growth conditions to the wood structure was researched on three species of conifers from various topoecological conditions in forest-steppe zone of Khakasia. By correlation analysis of climatic parameters (average temperatures and precipitation amounts on ten-day periods) of growth seasons and structure parameters of wood rings (tree ring width, maximum density, radial size of tracheids and thickness of cell walls) was revealed that at different conditions of growth significant transformation of climatic signal depending on the topoecological conditions and specific features of tree species takes place. It is shown also that the main target for external signal is cambial zone; subsequent processes of cells differentiation and their result are less depending on external conditions. Nevertheless, cells parameters and particularly their combination at trees from different topoecological conditions allow to receive additional and more detailed information about climatic features of growth season and to reveal “key” intervals of season, when climatic factors can be evaluated quantitatively.